Willow warblers prefer young, open, scrubby woodland with small trees, including human-altered habitats such as coppice and young plantations up to 10–20 years old. High amounts of birch, alder and willow, with good lichen amounts, and water features (e.g. streams), fields with large amounts of bracken and mosses, and patches of low bramble (for nest cover) are preferred, but it will use a wide range of other species, including young or open coniferous forests. Incorporating woodland ride edge thickets of varying structure and height is beneficial. They prefer damp woodland areas. Thicket forming shrubs like blackthorn provide pockets of habitat. Deer browsing can degrade the required low cover.
The highest population densities are found in Scandinavia (where it is the commonest bird of any), with up to 1,100 pairs per square kilometre, and a total population in Sweden and Finland of 24 million pairs. Lower densities occur further east, with peak densities of 27 pairs per square kilometre in central Siberia. Even lower densities are found on the southern edge of the breeding range, with just 9 pairs per square kilometre in Switzerland, and a total of just 100 pairs in the whole of northern Spain.
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