The great cormorant (Phalacrocorax carbo), known as the black shag or kawau in New Zealand, formerly also known as the great black cormorant across the Northern Hemisphere, the black cormorant in Australia, and the large cormorant in India, is a widespread member of the cormorant family of seabirds. The genus name is Latinised Ancient Greek, from φαλακρός (phalakros, "bald") and κόραξ (korax, "raven"), and carbo is Latin for "charcoal".
It breeds in much of the Old World, Australia, and the Atlantic coast of North America.
The 80–100 cm (30–40 in) long white-breasted cormorant P. c. lucidus, found in sub-Saharan Africa, has a white neck and breast. It is often treated as a full species, Phalacrocorax lucidus (e.g. Sibley & Monroe 1990, Sinclair, Hockey & Tarboton 2002).
In addition to the Australasian and African forms, other geographically distinct subspecies are recognised, including P. c. sinensis (western Europe to east Asia), P. c. maroccanus (north-western Africa), and P. c. hanedae (Japan).
Some authors treat all these as allospecies of a P. carbo superspecies group.
In New Zealand, the subspecies P. c. novaehollandiae is known as the black shag or by its Māori name: "kawau". The syntype is in the collection of the Museum of New Zealand Te Papa Tongarewa.
The great cormorant is a large black bird, but there is a wide variation in size in the species' wide range. Weight is reported to vary from 1.5 kg (3 lb 5 oz) to 5.3 kg (11 lb 11 oz).[8] Males are typically larger and heavier than females, with the nominate race (P. c. carbo) averaging about 10% larger in linear measurements than the smallest race in Europe (P. c. sinensis). The lightest average weights cited are in Germany (P. c. sinensis), where 36 males averaged 2.28 kg (5 lb 1⁄2 oz) and 17 females averaged 1.94 kg (4 lb 4+1⁄2 oz).[10] The highest come from Prince Edward Island in Canada (P. c. carbo), where 11 males averaged 3.68 kg (8 lb 2 oz) and 11 females averaged 2.94 kg (6 lb 7+1⁄2 oz). Length can vary from 70 to 102 cm (27+1⁄2 to 40 in) and wingspan from 121 to 160 cm (47+1⁄2 to 63 in). They are tied as the second largest extant species of cormorant after the flightless cormorant, with the Japanese cormorant averaging at a similar size. In bulk if not in linear dimensions, the blue-eyed shag species complex of the Southern Oceans are scarcely smaller at average.[10] It has a longish tail and yellow throat-patch. Adults have white patches on the thighs and on the throat in the breeding season. In European waters it can be distinguished from the common shag by its larger size, heavier build, thicker bill, lack of a crest and plumage without any green tinge. In eastern North America, it is similarly larger and bulkier than double-crested cormorant, and the latter species has more yellow on the throat and bill and lack the white thigh patches frequently seen on great cormorants. Great cormorants are mostly silent, but they make various guttural noises at their breeding colonies.
This is a very common and widespread bird species. It feeds on the sea, in estuaries, and on freshwater lakes and rivers. Northern birds migrate south and winter along any coast that is well-supplied with fish.
In Serbia, the cormorant lives in Vojvodina. However, after 1945 many artificial lakes were formed in Serbia; some of them became potential habitats for cormorants. Currently, on the Lake Ćelije, formed in 1980, there is a resident colony of cormorants, who nest there and are present throughout the year, except January–February 1985 and February 2012 when the lake surface was completely frozen.
The type subspecies, P. c. carbo, is found mainly in Atlantic waters and nearby inland areas: on western European coasts and east across the Palearctic to Siberia and to North Africa, the Faroe Islands, Iceland and Greenland; and on the eastern seaboard of North America. The subspecies P. c. novaehollandiae is found in Australian waters.
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