15-11-1993 AMBOSELI, KENYA - SOUTHERN LION (FEMALE) (Panthera leo ssp. melanochaita)

Panthera leo melanochaita is a lion subspecies in Southern and East Africa. In this part of Africa, lion populations are regionally extinct in Lesotho, Djibouti and Eritrea, and are threatened by loss of habitat and prey base, killing by local people in retaliation for loss of livestock, and in several countries also by trophy hunting. Since the turn of the 21st century, lion populations in intensively managed protected areas in Botswana, Namibia, South Africa and Zimbabwe have increased, but declined in East African range countries. In 2005, a Lion Conservation Strategy was developed for East and Southern Africa.

Results of a phylogeographic study indicate that lion populations in southern and eastern Africa form a major clade distinct from lion populations in West Africa, Central Africa and Asia. In 2017, the Cat Classification Task Force of the IUCN Cat Specialist Group subsumed lion populations according to the major clades into two subspecies, namely P. l. leo and P. l. melanochaita. Within P. l. melanochaita three subclades are clearly distinguishable. One from northeastern Africa, another one from southwestern Africa and a third one from southeastern Africa.

The type specimen for P. l. melanochaita was a black-maned lion from the Cape of Good Hope, known as the Cape lion. Phylogeographic analysis of lion samples from Gabon and the Republic of the Congo indicate their close genetic relation to P. l. melanochaita samples from Namibia and Botswana. It has been referred to as the Southern lion, Southern African lion, East-Southern African lion and the "southern subspecies".

15-11-1993 AMBOSELI, KENYA - SOUTHERN LION (FEMALE) (Panthera leo ssp. melanochaita)

The Southern lion is a lion subspecies from Southern and East Africa. In this region, lion populations were referred to by several regional names, including Katanga lion, Transvaal lion, Kalahari lion, Southeast African lion, Southwest African lion, Masai lion, Serengeti lion, Tsavo lion, and Uganda lion. It has also been referred to as 'Eastern-Southern African lion', 'Southern lion', and as 'southern subspecies'.

The fur of Southern lions varies in color from light buff to dark brown. They have rounded ears and a black tail tuft. Mane color varies from sandy, tawny, isabelline, and light reddish yellow to dark brown and black. Mane length varies from short to extending to knee joints and under the belly. Lions without a mane were observed in the Tsavo area. Mane development is related to age: older males have more extensive manes than younger ones; manes continue to grow up to the age of four to five years, long after lions become reproductively mature. 

Males living in the Kenyan highlands develop heavier manes than lions in the more humid and warmer lowlands of eastern and northern Kenya. White lions have occasionally been encountered in and around South Africa's Kruger National Park and the adjacent Timbavati Private Game Reserve. Their whitish fur is a rare morph caused by a double recessive allele. It has normal pigmentation in the eyes and skin. They were removed from the wild in the 1970s, thus decreasing the white lion gene pool.

15-11-1993 AMBOSELI, KENYA - OLIVE BABOON (Papio anubi)

The olive baboon (Papio anubis), also called the Anubis baboon, is a member of the family Cercopithecidae Old World monkeys. The species is the most wide-ranging of all baboons, being native to 25 countries throughout Africa, extending from Mali eastward to Ethiopia and Tanzania. Isolated populations are also present in some mountainous regions of the Sahara. It inhabits savannahs, steppes, and forests. The common name is derived from its coat colour, which is a shade of green-grey at a distance. A variety of communications, vocal and non-vocal, facilitate a complex social structure.

The olive baboon is named for its coat, which, at a distance, is a shade of green-grey. At closer range, its coat is multicoloured, due to rings of yellow-brown and black on the hairs. The hair on the baboon's face is coarser and ranges from dark grey to black. This coloration is shared by both sexes, although males have a mane of longer hair that tapers down to ordinary length along the back.

Besides the mane, the male olive baboon differs from the female in terms of weight, body and canine tooth size; males are, on average, 70 cm (28 in) tall while standing and females measure 60 cm (24 in) in height. The olive baboon is one of the largest species of monkey; along with chacma baboons, mandrills, and muriquis. The head-and-body length can range from 50 to 114 cm (20 to 45 in), with a species average of around 85 cm (33 in). At the shoulder on all fours, females average 55 cm (22 in) against males, which average 70 cm (28 in). The typical weight range for both sexes is reportedly 10–37 kg (22–82 lb), with males averaging 24 kg (53 lb) and females averaging 14.7 kg (32 lb). Some males may weigh as much as 50 kg (110 lb).

Like other baboons, the olive baboon has an elongated, dog-like muzzle. Its 38 to 58 cm (15 to 23 in) long tail and four-legged gait can make it seem canine. The tail almost looks as if it is broken, as it is erect for the first quarter, after which it drops down sharply. The bare patch of a baboon's rump is smaller in the olive baboon than in the Hamadryas baboon or Guinea baboon. The olive baboon, like most cercopithecines, has a cheek pouch with which to store food.

The species inhabits a strip of 25 equatorial African countries, very nearly ranging from the east to west coasts of the continent. The exact boundaries of this strip are not clearly defined, as the species' territory overlaps with that of other baboon species. In many places, this has resulted in cross-breeding between species. For example, considerable hybridisation has occurred between the olive baboon and the hamadryas baboon in Ethiopia. Cross-breeding with the yellow baboon and the Guinea baboon has also been observed. Although this has been noted, the hybrids have not as yet been well studied.

Throughout its wide range, the olive baboon can be found in a number of different habitats. It is usually classified as savannah-dwelling, living in the wide plains of the grasslands. The grasslands, especially those near open woodland, do make up a large part of its habitat, but the baboon also inhabits rainforests and deserts. Uganda and the Democratic Republic of the Congo, for instance, both support olive baboon populations in dense tropical forests.

15-11-1993 AMBOSELI, KENYA - EASTERN BLACK RHINOCEROS (Diceros bicornis ssp. michaeli)

The eastern black rhinoceros (Diceros bicornis michaeli), also known as the East African black rhinoceros, is a subspecies of the black rhinoceros. Its numbers are very low due to poaching for its horn, and it is listed as critically endangered.

The eastern black rhino is distinguishable from the southern subspecies as it has a longer, leaner, and more curved horn. Its skin is also very grooved. Diceros bicornis michaeli is also reportedly more aggressive than the other three subspecies of black rhino. They are browsers and are usually found in highland forest and savanna habitat.

Once located in Ethiopia, Somalia, Tanzania and Kenya, as of 2017 they can only be found in Kenya (594 animals), Rwanda and in northern Tanzania (80 animals). A population of currently 60 animals is kept outside its natural range in South Africa (Addo Elephant National Park). The population has declined 90% in the last three generations. In 2010 their total numbers were estimated at 740 animals, with an increasing trend. They are threatened mainly from illegal poaching for their horns.

The IUCN figures for Diceros bicornis michaeli also include those for black rhinos from South Sudan, Uganda, southwestern Ethiopia and western Kenya. These are referred to a separate subspecies (Diceros bicornis ladoensis) by some authorities. As the black rhinoceros population is extirpated in most of these areas, the status of the latter subspecies is unclear. Some animals of the Kenyan population may belong to it.

These black rhinos were examined by Benson and others through the Kenya Wildlife Service and had an article published by the African Journal of Ecology (Benson, 1, 791). Each individual rhino's relationship concerning “density dependence...sex ratio, and underlying growth rates” was the team's main focus. The rhino began increasing its population growth to the point when growth rate was nonexistent.

15-11-1993 AMBOSELI, KENYA - AFRICAN ELEPHANT (Loxodonta africana)

The African bush elephant (Loxodonta africana) is one of two living African elephant species. It is the largest living terrestrial animal and is distributed across 37 African countries. Since 2021, it has been listed as Endangered on the IUCN Red List. It is threatened foremost by habitat destruction, and in parts of its range also by poaching for meat and ivory.

The African bush elephant has grey skin with scanty hairs. Its large ears cover the whole shoulder and can grow as large as 2 m × 1.5 m (6 ft 7 in × 4 ft 11 in). Its large ears help to reduce body heat; flapping them creates air currents and exposes large blood vessels on the inner sides to increase heat loss during hot weather. The African bush elephant's ears are pointed and triangular-shaped. Its occipital plane slopes forward. Its back is shaped markedly concave. Its sturdy tusks are curved out and point forward. The trunk is a prehensile extension of the upper lip and nose. Short tactile hair grows on the trunk, which has two finger-like processes on the tip. This highly sensitive organ is innervated primarily by the trigeminal nerve and is thought to be manipulated by about 40,000-60,000 muscles. Because of this muscular structure, the trunk is so strong that elephants can use it for lifting about 3% of their own body weight. They use it for smelling, touching, feeding, drinking, dusting, sound production, loading, defending, and attacking. Both sexes have tusks, which erupt when they are 1-3 years old and grow throughout life. Tusks grow from deciduous teeth known as tushes that develop in the upper jaw and consist of a crown, root, and pulpal cavity, which are completely formed soon after birth. Tushes reach a length of 5 cm (2.0 in). The tusks of males grow faster than the tusks of females.

African bush elephants occur in Sub-Saharan Africa including Uganda, Kenya, Tanzania, Botswana, Zimbabwe, Namibia, Zambia, and Angola. They move between a variety of habitats, including forests, dry and seasonally flooded grasslands, woodlands, shrubland, and wetlands to mountain slopes. In Mali and Namibia, they also inhabit desert and semi-desert areas.

Savanna elephants live in a rather complex social hierarchy. These animals gather into family units, consisting of about 10 females and their offspring. Reaching maturity, male calves usually leave the family unit, forming bachelor herds or living solitarily. As a general rule, males socialize with these family groups only when mating. Meanwhile, several family herds may gather together, making up a 'clan'. Each clan is dominated by a female matriarch and can consist of as many as several hundred elephants. African bush elephants are very careful and protective animals. Allomothering is a common practice in this species: females can raise calves of other females of their herd. They protect and care for the calves of the herd, while all adults are sleeping. If a calf strays too far, these allomothers are responsible for retrieving the baby. These active animals are constantly on the move. These elephants forage during the daytime hours, wandering the home range of their herd. Savanna elephants freely communicate with conspecifics both verbally and non-verbally.

15-11-1993 AMBOSELI, KENYA - SOUTHERN LION (Panthera leo ssp. melanochaita)

 

Panthera leo melanochaita is a lion subspecies in Southern and East Africa. In this part of Africa, lion populations are regionally extinct in Lesotho, Djibouti and Eritrea, and are threatened by loss of habitat and prey base, killing by local people in retaliation for loss of livestock, and in several countries also by trophy hunting. Since the turn of the 21st century, lion populations in intensively managed protected areas in Botswana, Namibia, South Africa and Zimbabwe have increased, but declined in East African range countries. In 2005, a Lion Conservation Strategy was developed for East and Southern Africa.

Results of a phylogeographic study indicate that lion populations in southern and eastern Africa are forming a major clade distinct from lion populations in West Africa, Central Africa and Asia.[5] In 2017, the Cat Classification Task Force of the IUCN Cat Specialist Group subsumed lion populations according to the major clades into two subspecies, namely P. l. leo and P. l. melanochaita.[1] Within P. l. melanochaita three subclades are clearly distinguishable. One from northeastern Africa, another one from southwestern Africa and a third one from southeastern Africa.

The type specimen for P. l. melanochaita was a black-maned lion from the Cape of Good Hope, known as the Cape lion. Phylogeographic analysis of lion samples from Gabon and the Republic of the Congo indicate their close genetic relation to P. l. melanochaita samples from Namibia and Botswana. It has been referred to as the Southern lion, Southern African lion, East-Southern African lion and the "southern subspecies".

15-11-1993 AMBOSELI, KENYA - GRANT'S GAZELLE (MALE) (Nanger granti)

Grant's gazelle (Nanger granti) is a relatively large species of gazelle antelope, distributed from northern Tanzania to South Sudan and Ethiopia, and from the Kenyan coast to Lake Victoria. Its Swahili name is swala granti. It was named for a 19th-century British explorer, James Grant.

Grant's gazelle is genetically related to Soemmerring's gazelle (N. soemmerringii) and Thomson's gazelle (Eudorcas thomsonii) with Soemmering's gazelle being the closer relative. Grant's gazelle shows high genetic variation among its populations, although there is no geographic isolation. The differentiation of the species may have evolved during repeated expansion and contraction of arid habitats during the late Pleistocene era in which populations were possibly isolated. Grant's gazelle was formerly considered a member of the genus Gazella within the subgenus Nanger before Nanger was elevated to genus status. In 2021, the American Society of Mammalogists granted full species status to Bright's gazelle (Nanger notatus) and the Peter's gazelle (Nanger petersii).

15-11-1993 AMBOSELI, KENYA - COMMON WATERBUCK (MALE) (Kobus ellipsiprymnus ssp. ellipsiprymnus)

The waterbuck (Kobus ellipsiprymnus) is a large antelope found widely in sub-Saharan Africa. It is placed in the genus Kobus of the family Bovidae. It was first described by Irish naturalist William Ogilby in 1833. Its 13 subspecies are grouped under two varieties: the common or ellipsiprymnus waterbuck and the defassa waterbuck. The head-and-body length is typically between 177 and 235 cm (70 and 93 in) and the typical height is between 120 and 136 cm (47 and 54 in). In this sexually dimorphic antelope, males are taller and heavier than females. Males reach roughly 127 cm (50 in) at the shoulder, while females reach 119 cm (47 in). Males typically weigh 198–262 kg (437–578 lb) and females 161–214 kg (355–472 lb). Their coat colour varies from brown to grey. The long, spiral horns, present only on males, curve backward, then forward, and are 55–99 cm (22–39 in) long.

Waterbucks are rather sedentary in nature. As gregarious animals, they may form herds consisting of six to 30 individuals. These groups are either nursery herds with females and their offspring or bachelor herds. Males start showing territorial behaviour from the age of 5 years, but are most dominant from the six to nine. The waterbuck cannot tolerate dehydration in hot weather, and thus inhabits areas close to sources of water. Predominantly a grazer, the waterbuck is mostly found on grassland. In equatorial regions, breeding takes place throughout the year, but births are at their peak in the rainy season. The gestational period lasts 7–8 months, followed by the birth of a single calf.

15-11-1993 AMBOSELI, KENYA - SOUTHERN LION (FEMALE) (Panthera leo ssp. melanochaita)

Panthera leo melanochaita is a lion subspecies in Southern and East Africa. In this part of Africa, lion populations are regionally extinct in Lesotho, Djibouti and Eritrea, and are threatened by loss of habitat and prey base, killing by local people in retaliation for loss of livestock, and in several countries also by trophy hunting. Since the turn of the 21st century, lion populations in intensively managed protected areas in Botswana, Namibia, South Africa and Zimbabwe have increased, but declined in East African range countries. In 2005, a Lion Conservation Strategy was developed for East and Southern Africa.

Results of a phylogeographic study indicate that lion populations in southern and eastern Africa form a major clade distinct from lion populations in West Africa, Central Africa and Asia. In 2017, the Cat Classification Task Force of the IUCN Cat Specialist Group subsumed lion populations according to the major clades into two subspecies, namely P. l. leo and P. l. melanochaita. Within P. l. melanochaita three subclades are clearly distinguishable. One from northeastern Africa, another one from southwestern Africa and a third one from southeastern Africa.

15-11-1993 AMBOSELI, KENYA - SOUTHERN LION (AT KILL) (Panthera leo ssp. melanochaita)

Panthera leo melanochaita is a lion subspecies in Southern and East Africa. In this part of Africa, lion populations are regionally extinct in Lesotho, Djibouti and Eritrea, and are threatened by loss of habitat and prey base, killing by local people in retaliation for loss of livestock, and in several countries also by trophy hunting. Since the turn of the 21st century, lion populations in intensively managed protected areas in Botswana, Namibia, South Africa and Zimbabwe have increased, but declined in East African range countries. In 2005, a Lion Conservation Strategy was developed for East and Southern Africa.

Results of a phylogeographic study indicate that lion populations in southern and eastern Africa form a major clade distinct from lion populations in West Africa, Central Africa and Asia. In 2017, the Cat Classification Task Force of the IUCN Cat Specialist Group subsumed lion populations according to the major clades into two subspecies, namely P. l. leo and P. l. melanochaita. Within P. l. melanochaita three subclades are clearly distinguishable. One from northeastern Africa, another one from southwestern Africa and a third one from southeastern Africa.

The type specimen for P. l. melanochaita was a black-maned lion from the Cape of Good Hope, known as the Cape lion. Phylogeographic analysis of lion samples from Gabon and the Republic of the Congo indicate their close genetic relation to P. l. melanochaita samples from Namibia and Botswana. It has been referred to as the Southern lion, Southern African lion, East-Southern African lion and the "southern subspecies".

The Serengeti and Maasai Mara National Parks and Selous Game Reserve are lion strongholds in East Africa with stable lion populations

In East and Southern Africa, lion populations declined in:

In Ethiopia, where lion populations declined since at least the early 20th century due to trophy hunting by Europeans, killing of lions by local people out of fear, for illegal sale of skins and during civil wars. As of 2009, between seven and 23 lions were estimated to live in Nechisar National Park located in the Ethiopian Highlands. This small protected area is encroached by local people and their livestock. In 2012, lions were documented in cloud forest habitat of Kafa Biosphere Reserve.

Somalia since the early 20th century. Intensive poaching since the 1980s and civil unrest posed a threat to lion persistence.

Uganda to near extinction in the 20th century. In 2010, the lion population in Uganda was estimated at 408 ± 46 individuals in three protected areas including Queen Elizabeth, Murchison Falls and Kidepo Valley National Parks. Other protected areas in the country probably host less than 10 lions. Lions in Queen Elizabeth National Park form a contiguous population with lions in Virunga National Park.

Kenya in the 1990s due to poisoning of lions and poaching of lion prey species. At least 108 lions were killed between 2001 and 2006 in the Amboseli−Tsavo West−Tsavo East National Park network. As of 2006, there were an estimated 675 lions in the Tsavo national parks, out of the 2,000 total in Kenya. Between 2004 and 2013, lion guardians around Amboseli National Park identified 65 lions in an area of 3,684 km2 (1,422 sq mi). Lion populations in Kenya and Tanzania are fragmented over 17 patches ranging in size from 86 to 127,515 km2 (33 to 49,234 sq mi).

Rwanda and Tanzania due to killing of lions during the Rwandan Civil War and ensuing refugee crisis in the 1990s. A small population was present in Rwanda's Akagera National Park, estimated at 35 individuals at most in 2004. Lions were reintroduced to this national park in 2015.

Malawi and Zambia due to illegal hunting of prey species in protected areas.

Botswana due to intensive hunting and conversion of natural habitats for settlements since the early 19th century. In Northern Tuli Game Reserve, 19 lions died between 2005 and 2011 due to poaching, trophy hunting and snaring.

Namibia due to massive killing of lions by farmers since at least the 1970s. In 2010, the small and isolated lion population in the Kalahari was estimated at 683 to 1,397 individuals in three protected areas, the Kgalagadi Transfrontier Park, the Kalahari Gemsbok and Gemsbok National Parks.

South Africa since the early 19th century in the Natal and Cape Provinces south of the Orange River, where the Cape lion population was eradicated by 1860. A few decades later, lions in the Highveld north of the Orange River were also eradicated. In Transvaal, lions occurred historically in the Highveld as well, but were restricted to eastern Transvaal's Bushveld by the 1970s. Between 2000 and 2004, 34 lions were reintroduced to eight protected areas in the Eastern Cape Province, including Addo Elephant National Park. In Venetia Limpopo Nature Reserve, 18 lions were trophy hunted and 11 euthanized between 2005 and 2011.

Contemporary lion distribution and habitat quality in East and Southern Africa was assessed in 2005, and Lion Conservation Units (LCU) mapped. Between 2002 and 2012, educated guesses for size of populations in these LCUs ranged from 33,967 to 32,000 individuals. The LCUs Ruaha−Rungwa, Serengeti−Mara, Tsavo−Mkomazi and Selous in East Africa, as well as Luangwa, Kgalagadi, Okavango−Hwange, Mid−Zambezi, Niassa and Greater Limpopo in Southern Africa are currently considered lion strongholds. These LCUs host more than 500 individuals each, and the population trend is stable there as of 2012.

14-11-1993 AMBOSELI, KENYA - NILE MONITOR (Varanus niloticus)

The Nile monitor (Varanus niloticus) is a large member of the monitor family (Varanidae) found throughout most of Sub-Saharan Africa, particularly in drier regions, and along the Nile River and its tributaries in East Africa. Additionally, there are modern, invasive populations in North America. The population found in West African forests and savannahs is sometimes recognized as a separate species, the West African Nile monitor (V. stellatus). While it is dwarfed by its larger relatives, such as the Komodo dragon, the Asian water monitor or the crocodile monitor, it is still one of the largest lizards in the world, reaching (and even surpassing) Australia’s perentie in size. Other common names include the African small-grain lizard, as well as iguana and various forms derived from it, such as guana, water leguaan or river leguaan (leguan, leguaan, and likkewaan mean monitor lizard in South African English, and can be used interchangeably).

A feral population of Nile monitors (descended from escaped or intentionally-released pets) has become established in several locations in South Florida.[8] In addition to any illegally-released animals, it is speculated that during particularly intense hurricane seasons in Florida, many reptiles potentially escape when their enclosures are damaged or inadvertently unlocked; as Florida has a semi-tropical to tropical climate, many reptiles are housed outdoors, and poorly-secured enclosures may become damaged during bad storms. Along with Nile monitors, Florida is infamous for its feral populations of agamas, Argentine black and white tegus, Burmese pythons, green iguanas, Madagascar giant day geckos, and panther and veiled chameleons, among others. Many of these species are thought to be descendants of hurricane escapees.

10-11-1993 MOUNTAIN LODGE, KENYA - VERVET MONKEY (Chlorocebus pygerythrus)

The vervet monkey (Chlorocebus pygerythrus), or simply vervet, is an Old World monkey of the family Cercopithecidae native to Africa. The term "vervet" is also used to refer to all the members of the genus Chlorocebus. The five distinct subspecies can be found mostly throughout Southern Africa, as well as some of the eastern countries. These mostly herbivorous monkeys have black faces and grey body hair color, ranging in body length from about 40 cm (16 in) for females, to about 50 cm (20 in) for males.

In addition to behavioral research on natural populations, vervet monkeys serve as a nonhuman primate model for understanding genetic and social behaviors of humans. They have been noted for having human-like characteristics, such as hypertension, anxiety, and social and dependent alcohol use. Vervets live in social groups ranging from 10 to 70 individuals, with males moving to other groups at the time of sexual maturity. Studies done on vervet monkeys involve their communication and alarm calls, specifically in regard to kin and group recognition, and particular predator sightings.

The vervet monkey ranges throughout much of Southern and East Africa, being found from Ethiopia and extreme southern South Sudan all the way down to South Africa. It is not found west of the East African Rift or the Luangwa River, where it is replaced by the closely related malbrouck (C. cynosuros) species. The vervet monkey inhabits savanna, riverine woodland, coastal forest, and mountains up to 4000 m (13,100 ft). They are adaptable and able to persist in secondary and/or highly fragmented vegetation, including cultivated areas, and sometimes are found living in both rural and urban environments. Annual home range size has been observed to be as high as 176 ha, with an average population density of 54.68 animals/km2.

Vervet monkeys that are naturalised (introduced by humans) are found in Cape Verde, Ascension Island, Saint Helena, St Kitts and Nevis, and Barbados. Dania Beach, Florida, is home to about 40 introduced vervets.

The vervet monkey very much resembles a gray langur, having a black face with a white fringe of hair, while its overall hair color is mostly grizzled-grey. The species exhibits sexual dimorphism; the males are larger in weight and body length and may be recognized by a turquoise-blue scrotum. Adult males weigh between 3.9 and 8.0 kg (8.6 and 17.6 lb), averaging 5.5 kg (12 lb), and have a body length between 420 and 600 mm (17 and 24 in), averaging 490 mm (19 in) from the top of the head to the base of the tail. Adult females weigh between 3.4 and 5.3 kg (7.5 and 11.7 lb) and average 4.1 kg (9.0 lb), and measure between 300 and 495 mm (11.8 and 19.5 in), averaging 426 mm (16.8 in).

When males reach sexual maturity, they move to a neighboring group. Often, males move with a brother or peer, presumably for protection against aggression by males and females of the resident group. Groups that had previously transferred males show significantly less aggression upon the arrival of another male. In almost every case, males migrate to adjacent groups. This obviously increases benefits in regard to distance traveled, but also reduces the amount of genetic variance, increasing the likelihood of inbreeding.

Females remain in their groups throughout life. Separate dominance hierarchies are found for each sex. Male hierarchies are determined by age, tenure in the group, fighting abilities, and allies, while female hierarchies are dependent on maternal social status. A large proportion of interactions occurs between individuals that are similarly ranked and closely related. Between unrelated individuals, female competition exists for grooming members of high-ranking families, presumably to gain more access to resources. These observations suggest individual recognition is possible and enables discrimination of genetic relatedness and social status. Interactions between different groups are variable, ranging from highly aggressive to friendly. Furthermore, individuals seem to be able to recognise cross-group vocalisations, and identify from and to which monkey each call is intended, even if the call is made by a subadult male, which is likely to transfer groups. This suggests the members within a group are actively monitoring the activity of other groups, including the movement of individuals within a group.

Within groups, aggression is directed primarily at individuals that are lower on the hierarchy. Once an individual is three years or older, it is considerably more likely to be involved in conflict. Conflict often arises when one group member shows aggression toward a close relative of another. Further, both males and females may redirect aggression towards individuals in which both had close relatives that were previously involved in a conflict. This suggests complex recognition not only of individuals, but also of associations between individuals. This does not suggest recognition of other's individual kinship bonds is possible, but rather that discrimination of social relationships does occur.

14-11-1993 MOUNTAIN LODGE, KENYA - DEFASSA WATERBUCK (FEMALE) (Kobus ellipsiprymnus ssp. defassa)

The waterbuck (Kobus ellipsiprymnus) is a large antelope found widely in sub-Saharan Africa. It is in the genus Kobus of the family Bovidae. It was first described by Irish naturalist William Ogilby in 1833. Its 13 subspecies are grouped under two varieties: the common or ellipsiprymnus waterbuck and the defassa waterbuck. Their coat colour varies from brown to grey. The long, spiral horns, present only on males, curve backward, then forward, and are 55–99 cm (22–39 in) long.

Waterbucks are rather sedentary in nature. As gregarious animals, they may form herds consisting of six to thirty individuals. These groups are either nursery herds with females and their offspring or bachelor herds. Males start showing territorial behaviour from the age of five years, but are most dominant from six to nine. The waterbuck cannot tolerate dehydration in hot weather, and thus inhabits areas close to sources of water. Predominantly a grazer, the waterbuck is mostly found on grassland. In equatorial regions, breeding takes place throughout the year, but births are at their peak in the rainy season. The gestational period lasts 7–8 months, followed by the birth of a single calf.

Waterbucks inhabit scrub and savanna areas along rivers, lakes, and valleys. Due to their requirement for grasslands and water, waterbucks have a sparse ecotone distribution. The IUCN lists the waterbuck as being of least concern. More specifically, the common waterbuck is listed as of least concern, while the defassa waterbuck is near threatened. The population trend for both is downwards, especially that of the defassa, with large populations being eliminated from certain habitats because of poaching and human disturbance.

14-11-1993 AMBOSELI, KENYA - BUSH HYRAX (Heterohyrax brucei)

The yellow-spotted rock hyrax or bush hyrax (Heterohyrax brucei) is a species of mammal in the family Procaviidae. It is found in Angola, Botswana, Burundi, the Democratic Republic of the Congo, southern Egypt, Eritrea, Ethiopia, Kenya, Malawi, Mozambique, Rwanda, Somalia, northern South Africa, South Sudan, Sudan, Tanzania, Uganda, Zambia, and Zimbabwe. Its natural habitats are dry savanna and rocky areas. Hyrax comes from the Greek word ὕραξ, or shrew-mouse.

Heterohyrax is a bush hyrax, as opposed to a rock hyrax (Procavia capensis) or a tree hyrax (Dendrohyrax). Although difficult to distinguish in the field, the bush hyrax differs from the rock hyrax in being smaller and less heavily built and having a narrower muzzle. 

Common names for the bush hyrax include yellow-spotted hyrax, bush hyrax, hogger hyrax, yellow-spotted rock hyrax, and daman de steppe (French).

The genus Heterohyrax contains one extinct species, H. auricampensis, and one living, H. brucei. Within H. brucei are 25 recognized subspecies.

14-11-1993 MOUNTAIN LODGE, KENYA - COMMON WATERBUCK (FEMALE) (Kobus ellipsiprymnus ssp. ellipsiprymnus)

The waterbuck (Kobus ellipsiprymnus) is a large antelope found widely in sub-Saharan Africa. It is in the genus Kobus of the family Bovidae. It was first described by Irish naturalist William Ogilby in 1833. Its 13 subspecies are grouped under two varieties: the common or ellipsiprymnus waterbuck and the defassa waterbuck. Their coat colour varies from brown to grey. The long, spiral horns, present only on males, curve backward, then forward, and are 55–99 cm (22–39 in) long.

Waterbucks are rather sedentary in nature. As gregarious animals, they may form herds consisting of six to thirty individuals. These groups are either nursery herds with females and their offspring or bachelor herds. Males start showing territorial behaviour from the age of five years, but are most dominant from six to nine. The waterbuck cannot tolerate dehydration in hot weather, and thus inhabits areas close to sources of water. Predominantly a grazer, the waterbuck is mostly found on grassland. In equatorial regions, breeding takes place throughout the year, but births are at their peak in the rainy season. The gestational period lasts 7–8 months, followed by the birth of a single calf.

14-11-1993 MOUNTAIN LODGE, KENYA - AFRICAN SAVANNAH ELEPHANT

The African bush elephant (Loxodonta africana), also known as the African savanna elephant, is a species of elephant native to sub-Saharan Africa. It is one of three extant elephant species and, along with the African forest elephant, one of two extant species of African elephant. It is the largest living terrestrial animal, with fully grown bulls reaching an average shoulder height of 3.04–3.36 metres (10.0–11.0 ft) and a body mass of 5.2–6.9 tonnes (11,000–15,000 lb); the largest recorded specimen had a shoulder height of 3.96 metres (13.0 ft) and an estimated body mass of 10.4 tonnes (23,000 lb). The African bush elephant is characterised by its long prehensile trunk with two finger-like processes; a convex back; large ears which help reduce body heat; and sturdy tusks that are noticeably curved. The skin is grey with scanty hairs, and bending cracks which support thermoregulation by retaining water.

The African bush elephant inhabits a variety of habitats such as forests, grasslands, woodlands, wetlands and agricultural land. It is a mixed herbivore feeding mostly on grasses, creepers, herbs, leaves, and bark. The average adult consumes about 150 kg (330 lb) of vegetation and 230 L (51 imp gal; 61 US gal) of water each day. A social animal, the African bush elephant often travels in herds composed of cows and their offspring. Adult bulls usually live alone or in small bachelor groups. During the mating season, males go through a process called musth; a period of high testosterone levels and heightened aggression. For females, the menstrual cycle lasts three to four months, and gestation around 22 months, the longest of any mammal.

14-11-1993 MOUNTAIN LODGE, KENYA - VON DER DECKEN'S HORNBILL (Tockus deckeni)

Von der Decken's hornbill (Tockus deckeni) is a hornbill found in East Africa, especially to the east of the East African Rift, from Ethiopia south to Tanzania. It is found mainly in thorn scrub and similar arid habitats. Jackson's hornbill is often treated as a subspecies of it. It was named after the German explorer Baron Karl Klaus von der Decken (1833–1865).[2]

This species is a small hornbill with blackish upperparts and mainly whitish underparts and head. It has a long tail and a long curved bill which lacks a casque. It is similar to the red-billed hornbill except for the bill colour and the lack of spotting on the wing coverts in both male and female.

The species shows sexual dimorphism; the female has a black bill, whereas the male has a red bill with a cream tip and a black cutting edge.

During incubation, the female lays two or three white eggs in a tree hole, which is blocked off with a cement made of mud, droppings and fruit pulp. There is only one narrow aperture, just big enough for the male to transfer food to the mother and chicks.

When the chicks and the female are too big for the nest, the mother breaks out and rebuilds the wall. Then both parents feed the chicks.

Captive breeding can be achieved by providing a small barrel or hollow tree with an entrance hole 5 inches (13 cm) tall and 3 inches (7.6 cm) wide. The birds will block off the hole once the hen is ready.

Von der Decken's hornbill is omnivorous, taking insects, fruit and seeds. It feeds mainly on the ground and will form flocks outside the breeding season. In captivity, Von Der Decken's hornbill will readily eat papaya, cantaloupe, blueberries, bananas, and apples. Live food such as crickets and mealworms should also be offered daily. Small rodents are readily taken but should only be offered two or three times a week.

14-11-1993 MOUNTAIN LODGE, KENYA - EASTERN PALM FORESTER BUTTERFLY (Bebearia orientis)

Bebearia orientis, the eastern palm forester, is a butterfly in the family Nymphalidae. It is found in Somalia, Kenya, Tanzania, Mozambique, Malawi, Zambia and Zimbabwe. The habitat consists of forests.

Adults are attracted to fermented fruit. They are probably on wing year round.

The larvae feed on Phoenix reclinata, Raphia farinifera and probably Cocos nucifera.

14-11-1993 MOUNTAIN LODGE, KENYA - OLIVE BABOON (Papio anubi)

The olive baboon (Papio anubis), also called the Anubis baboon, is a member of the family Cercopithecidae Old World monkeys. The species is the most wide-ranging of all baboons, being native to 25 countries throughout Africa, extending from Mali eastward to Ethiopia and Tanzania. Isolated populations are also present in some mountainous regions of the Sahara. It inhabits savannahs, steppes, and forests. The common name is derived from its coat colour, which is a shade of green-grey at a distance. A variety of communications, vocal and non-vocal, facilitate a complex social structure.

The olive baboon is named for its coat, which, at a distance, is a shade of green-grey. At closer range, its coat is multicoloured, due to rings of yellow-brown and black on the hairs. The hair on the baboon's face is coarser and ranges from dark grey to black. This coloration is shared by both sexes, although males have a mane of longer hair that tapers down to ordinary length along the back.

14-11-1993 MOUNTAIN LODGE, KENYA - AFRICAN SAVANNA ELEPHANT (Loxodonta Africana)

The African bush elephant (Loxodonta africana), also known as the African savanna elephant, is a species of elephant native to sub-Saharan Africa. It is one of three extant elephant species and, along with the African forest elephant, one of two extant species of African elephant. It is the largest living terrestrial animal, with fully grown bulls reaching an average shoulder height of 3.04–3.36 metres (10.0–11.0 ft) and a body mass of 5.2–6.9 tonnes (5.7–7.6 short tons); the largest recorded specimen had a shoulder height of 3.96 metres (13.0 ft) and an estimated body mass of 10.4 tonnes (11.5 short tons). The African bush elephant is characterised by its long prehensile trunk with two finger-like processes; a convex back; large ears which help reduce body heat; and sturdy tusks that are noticeably curved. The skin is grey with scanty hairs, and bending cracks which support thermoregulation by retaining water.

The African bush elephant (Loxodonta africana), also known as the African savanna elephant, is a species of elephant native to sub-Saharan Africa. It is one of three extant elephant species and, along with the African forest elephant, one of two extant species of African elephant. It is the largest living terrestrial animal, with fully grown bulls reaching an average shoulder height of 3.04–3.36 metres (10.0–11.0 ft) and a body mass of 5.2–6.9 tonnes (5.7–7.6 short tons); the largest recorded specimen had a shoulder height of 3.96 metres (13.0 ft) and an estimated body mass of 10.4 tonnes (11.5 short tons). The African bush elephant is characterised by its long prehensile trunk with two finger-like processes; a convex back; large ears which help reduce body heat; and sturdy tusks that are noticeably curved. The skin is grey with scanty hairs, and bending cracks which support thermoregulation by retaining water.

Since 2021, the African bush elephant has been listed as Endangered on the IUCN Red List. It is threatened foremost by habitat destruction, and in parts of its range also by poaching for meat and ivory. Between 2003 and 2015, the illegal killing of 14,606 African bush elephants was reported by rangers across 29 range countries. Chad is a major transit country for smuggling of ivory in West Africa. This trend was curtailed by raising penalties for poaching and improving law enforcement. Poaching of the elephant has dated back to the 1970s and 80s, which were considered the largest killings in history. In human culture, elephants have been extensively featured in literature, folklore and media, and are most valued for their large tusks in many places.

10-11-1993 AMBOSELI, KENYA - BLACK BELLIED BUSTARD (Lissotis melanogaster)

The black-bellied bustard (Lissotis melanogaster), also known as the black-bellied korhaan, is an African ground-dwelling bird in the bustard family.

The black-bellied bustard is 58–65 cm (23–26 in) long. The bill and legs are dull yellow. The male's upperparts have black and brown marks on a tawny buff background; the underparts are black. The head is boldly patterned with black, white and buff. The neck, long and thin for a bustard, is buffy brown with a thin black line down the front that joins the black breast. The tail is brown and buff with four or five narrow dark brown bands. The upper surface of the wings is white with a brown triangle at the base; the flight feathers have black tips except for the outer secondary feathers. The white of the wings is visible when the bird stands, contrasting with the black underparts.

The female is plain buff, cryptically marked with darker brown mottling on the back and vermiculation (narrow wavy bands) on the neck and breast. The juvenile is duller and darker, with a dark grey crown and buff spots on the wing. The neck and rump patterns of both sexes, the male's white chin and lores, and the female's vermiculations are points that distinguish this species from its close relative, Hartlaub's bustard.

It is found in savanna, cultivated fields and tall open grassland in sub-Saharan Africa. This species is also found in the areas of S. Africa, Zimbabwe, Botswana and some areas of Tanzania and Nigeria. It prefers higher rainfall than Hartlaub's bustard and in many areas occurs only following heavy rain.

10-11-1993 MASAI MARA, KENYA - BLUE WILDEBEEST (Connochaetes taurinus ssp. taurinus)

The blue wildebeest (Connochaetes taurinus), also called the common wildebeest, white-bearded gnu or brindled gnu, is a large antelope and one of the two species of wildebeest. It is placed in the genus Connochaetes and family Bovidae, and has a close taxonomic relationship with the black wildebeest. The blue wildebeest is known to have five subspecies. This broad-shouldered antelope has a muscular, front-heavy appearance, with a distinctive, robust muzzle. Young blue wildebeest are born tawny brown, and begin to take on their adult coloration at the age of 2 months. The adults' hues range from a deep slate or bluish-gray to light gray or even grayish-brown. Both sexes possess a pair of large curved horns.

The blue wildebeest is an herbivore, feeding primarily on short grasses. It forms herds which move about in loose aggregations, the animals being fast runners and extremely wary. The mating season begins at the end of the rainy season and a single calf is usually born after a gestational period of about 8.5 months. The calf remains with its mother for 8 months, after which it joins a juvenile herd. Blue wildebeest are found in short-grass plains bordering bush-covered acacia savannas in southern and eastern Africa, thriving in areas that are neither too wet nor too arid. Three African populations of blue wildebeest take part in a long-distance migration, timed to coincide with the annual pattern of rainfall and grass growth on the short-grass plains where they can find the nutrient-rich forage necessary for lactation and calf growth.

The blue wildebeest is native to Angola, Botswana, Eswatini, Kenya, Mozambique, South Africa, Tanzania, Zambia, and Zimbabwe. Today, it is extinct in Malawi, but has been successfully reintroduced in Namibia. The southern limit of the blue wildebeest range is the Orange River, while the western limit is bounded by Lake Victoria and Mount Kenya. The blue wildebeest is widespread and is being introduced into private game farms, reserves, and conservancies. The International Union for Conservation of Nature and Natural Resources rates the blue wildebeest as being of least concern. The population has been estimated to be around 1.5 million, and the population trend is stable.

10-11-1993 MASAI MARA, KENYA - BLUE WILDEBEEST (Connochaetes taurinus ssp. taurinus)

The blue wildebeest is an herbivore, feeding primarily on short grasses. It forms herds which move about in loose aggregations, the animals being fast runners and extremely wary. The mating season begins at the end of the rainy season and a single calf is usually born after a gestational period of about 8.5 months. The calf remains with its mother for 8 months, after which it joins a juvenile herd. Blue wildebeest are found in short-grass plains bordering bush-covered acacia savannas in southern and eastern Africa, thriving in areas that are neither too wet nor too arid. Three African populations of blue wildebeest take part in a long-distance migration, timed to coincide with the annual pattern of rainfall and grass growth on the short-grass plains where they can find the nutrient-rich forage necessary for lactation and calf growth.

The blue wildebeest is native to Angola, Botswana, Eswatini, Kenya, Mozambique, South Africa, Tanzania, Zambia, and Zimbabwe. Today, it is extinct in Malawi, but has been successfully reintroduced in Namibia. The southern limit of the blue wildebeest range is the Orange River, while the western limit is bounded by Lake Victoria and Mount Kenya. The blue wildebeest is widespread and is being introduced into private game farms, reserves, and conservancies. The International Union for Conservation of Nature and Natural Resources rates the blue wildebeest as being of least concern. The population has been estimated to be around 1.5 million, and the population trend is stable.

10-11-1993 MASAI MARA, KENYA - WHITE BACKED VULTURE (Gyps africanus)

The White-backed Vulture (Gyps africanus) is a medium-sized vulture, characterized by down feathers on the head and neck, broad wings, and short tail feathers. It boasts a distinctive white neck ruff, with adults displaying a whitish back that contrasts starkly with the rest of their dark plumage. Juveniles present a more uniform dark appearance. This species typically weighs between 4.2 to 7.2 kg, measures 78 to 98 cm in length, and spans a wingspan of 1.96 to 2.25 m.

To identify the white-backed vulture, look for its white neck ruff and the contrasting white back against dark plumage in adults. Juveniles are darker overall. The species has a relatively short tail and broad wings suited for soaring.

The white-backed vulture is found in a variety of habitats across Africa, from the Sahel region to the savannahs of the south. It prefers wooded savannahs where it can find large trees suitable for nesting.

DistributionThis vulture is widespread across Africa, from Senegal and Gambia in the west, through Ethiopia and Somalia in the east, and down to Mozambique, Zimbabwe, Botswana, Namibia, and South Africa in the south.

White-backed vultures are social creatures, often seen circling the skies in search of carrion. They feed in groups and are typically among the first vultures to arrive at a carcass, asserting dominance over other species. After gorging themselves, they may rest with wings spread to bask in the sun.

White-backed vultures nest in trees, showing a preference for tall species such as Acacias. Their nests are large, constructed with sticks, and lined with leaves and grasses. The breeding cycle is lengthy, with an incubation period of around 8 weeks and a nestling period of 4-5 months.

10-11-1993 MASAI MARA, KENYA - COMMON WATERBUCK (MALE) (Kobus ellipsiprymnus ssp. ellipsiprymnus)

The waterbuck (Kobus ellipsiprymnus) is a large antelope found widely in sub-Saharan Africa. It is in the genus Kobus of the family Bovidae. It was first described by Irish naturalist William Ogilby in 1833. Its 13 subspecies are grouped under two varieties: the common or ellipsiprymnus waterbuck and the defassa waterbuck. Their coat colour varies from brown to grey. The long, spiral horns, present only on males, curve backward, then forward, and are 55–99 cm (22–39 in) long.

Waterbucks are rather sedentary in nature. As gregarious animals, they may form herds consisting of six to thirty individuals. These groups are either nursery herds with females and their offspring or bachelor herds. Males start showing territorial behaviour from the age of five years, but are most dominant from six to nine. The waterbuck cannot tolerate dehydration in hot weather, and thus inhabits areas close to sources of water. Predominantly a grazer, the waterbuck is mostly found on grassland. In equatorial regions, breeding takes place throughout the year, but births are at their peak in the rainy season. The gestational period lasts 7–8 months, followed by the birth of a single calf.

Waterbucks inhabit scrub and savanna areas along rivers, lakes, and valleys. Due to their requirement for grasslands and water, waterbucks have a sparse ecotone distribution. The IUCN lists the waterbuck as being of least concern. More specifically, the common waterbuck is listed as of least concern, while the defassa waterbuck is near threatened. The population trend for both is downwards, especially that of the defassa, with large populations being eliminated from certain habitats because of poaching and human disturbance.