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Thursday, 10 November 1994

10-11-1994 NOISE-BE, MADAGASCAR - BLACK LEMUR (Eulemur macaco)



The black lemur (Eulemur macaco) is a species of lemur from the family Lemuridae. Like all lemurs, it is endemic to Madagascar. Originally, the species was thought to have two subspecies, Eulemur macaco macaco and Eulemur macaco flavifrons, both of which were elevated to species status by Mittermeier et al. in 2008 to Eulemur macaco and Eulemur flavifrons respectively. The most startling difference between the two species is the eye colour; Eulemur flavifrons, the blue-eyed black lemur, has blue eyes, while Eulemur macaco, the black lemur, has brown or orange eyes, and also has ear tufts.


Both species live in northwest Madagascar. The black lemur occurs in moist forests in the Sambirano region of Madagascar and on nearby islands. The blue-eyed black lemur is restricted to the Sahamalaza Peninsula and adjacent forests. There are reports of the two species hybridizing where their ranges overlap in Manongarivo Special Reserve.


The black lemur is between 90 and 110 centimeters in length, of which 51–65 centimeters are tail. Weight typically ranges between 1.8 and 2.0 kilograms. The black lemur displays sexual dimorphism in coloration. Males have black or dark chocolate fur, while females' fur is of a lighter brown color, generally medium brown, chestnut brown or even orange-brown. Males have large black ear tufts, while females have large white ear tufts.


The only other Eulemur species that occurs within the range of the black lemur is the common brown lemur, which overlaps with the black lemur at the extreme southern and eastern edge of its range, and the red-bellied lemur, on the Tsaratanana Massif. The common brown lemur and the red-bellied lemur have different coloration and do not show the extreme sexual dimorphism of the black lemur, making confusion between the black lemur and the other species unlikely.

10-11-1994 NOISE-BE, MADAGASCAR - BLACK LEMUR (Eulemur macaco)


The black lemur (Eulemur macaco) is a species of lemur from the family Lemuridae. Like all lemurs, it is endemic to Madagascar. Originally, the species was thought to have two subspecies, Eulemur macaco macaco and Eulemur macaco flavifrons, both of which were elevated to species status by Mittermeier et al. in 2008 to Eulemur macaco and Eulemur flavifrons respectively. The most startling difference between the two species is the eye colour; Eulemur flavifrons, the blue-eyed black lemur, has blue eyes, while Eulemur macaco, the black lemur, has brown or orange eyes, and also has ear tufts.


Both species live in northwest Madagascar. The black lemur occurs in moist forests in the Sambirano region of Madagascar and on nearby islands. The blue-eyed black lemur is restricted to the Sahamalaza Peninsula and adjacent forests. There are reports of the two species hybridizing where their ranges overlap in Manongarivo Special Reserve.


The black lemur is between 90 and 110 centimeters in length, of which 51–65 centimeters are tail. Weight typically ranges between 1.8 and 2.0 kilograms. The black lemur displays sexual dimorphism in coloration. Males have black or dark chocolate fur, while females' fur is of a lighter brown color, generally medium brown, chestnut brown or even orange-brown. Males have large black ear tufts, while females have large white ear tufts.


The only other Eulemur species that occurs within the range of the black lemur is the common brown lemur, which overlaps with the black lemur at the extreme southern and eastern edge of its range, and the red-bellied lemur, on the Tsaratanana Massif. The common brown lemur and the red-bellied lemur have different coloration and do not show the extreme sexual dimorphism of the black lemur, making confusion between the black lemur and the other species unlikely.

Saturday, 5 November 1994

5-11-1994 KANDY, SRI LANKA - SRI LANKAN ELEPHANT (Elephas maximus ssp. maximus)

The Asian elephant (Elephas maximus), also known as the Asiatic elephant, is the only living Elephas species. It is the largest living land animal in Asia and the second largest living elephantid in the world. It is characterised by its long trunk with a single finger-like processing; large tusks in males; laterally folded large ears and wrinkled grey skin that is partly depigmented on the trunk, ears or neck. Adult males average 4 t (4.4 short tons) in weight and females 2.7 t (3.0 short tons). It has a large and well developed neocortex of the brain, is highly intelligent and self-aware being able to display behaviours associated with grief, learning and greeting. Three subspecies are recognised—E. m. maximus, E. m. indicus and E. m. sumatranus.

The Asian elephant is distributed in the Indian subcontinent and Southeast Asia, from India in the west to Borneo in the east, and Nepal in the north to Sumatra in the south. It frequently inhabits grasslands, tropical evergreen forests, semi-evergreen forests, moist deciduous forests, dry deciduous forests and dry thorn forests. It is herbivorous, eating about 150 kg (330 lb) of vegetation per day. Cows and calves form groups, while males remain solitary or form "bachelor groups" with other males. During the breeding season, males temporarily join female groups to mate. Wild Asian elephants live to be about 60 years old. While female captive elephants are recorded to have lived beyond 60 years when kept in semi-natural surroundings, Asian elephants die at a much younger age in captivity; captive populations are declining due to a low birth and high death rate.

Since 1986, the Asian elephant has been listed as Endangered on the IUCN Red List, as the population has declined by at least 50 per cent over the last three elephant generations, which is about 60–75 years. It is primarily threatened by loss of habitat, habitat degradation, fragmentation and poaching. The earliest indications of captive use of Asian elephants are engravings on seals of the Indus Valley civilisation dated to the 3rd millennium BC.

Monday, 8 August 1994

8-8-1994 JAIPUR, INDIA - NORTHERN PLAINS GREY LANGUR (Semnopithecus entellus)


The northern plains gray langur (Semnopithecus entellus), also known as the sacred langur, Bengal sacred langur and Hanuman langur, is a species of primate in the family Cercopithecidae.

The northern plains gray langur belongs to the genus Semnopithecus along with the other Indian langurs. The southern plains gray langur was once classified as a subspecies of S. entellus, i.e., S. entellus dussumieri and later regarded as a separate species, i.e., S. dussumieri, but is now regarded as an invalid taxon. Most of the specimens that had been regarded as Semnopithecus dussumieri fall within the revised range of Semnopithecus entellus.

The fur of adults is mostly light colored, with darker fur on the back and limbs, and the face, ears, hands and feet are all black. Infants are brown. Body size excluding tail ranges from 45.1 cm (17.8 in) to 78.4 cm (30.9 in) long, and the tail length is between 80.3 cm (31.6 in) and 111.8 cm (44.0 in). Adult males weigh between 16.9 kg (37 lb) and 19.5 kg (43 lb) while adult females weigh between 9.5 kg (21 lb) and 16.1 kg (35 lb).

The range of the northern plains gray langur covers a large portion of India south of the Himalayas south to the Tapti River and the Krishna River. It is thought to have been introduced to western Bangladesh by Hindu pilgrims on the bank of the Jalangi River.

Wednesday, 17 November 1993

17-11-1993 TSAVO NAT PARK, KENYA - SAVANNAH ELEPHANT (Loxodonta Africana)


The African bush elephant (Loxodonta africana), also known as the African savanna elephant, is a species of elephant native to sub-Saharan Africa. It is one of three extant elephant species and, along with the African forest elephant, one of two extant species of African elephant. It is the largest living terrestrial animal, with fully grown bulls reaching an average shoulder height of 3.04–3.36 metres (10.0–11.0 ft) and a body mass of 5.2–6.9 tonnes (11,000–15,000 lb); the largest recorded specimen had a shoulder height of 3.96 metres (13.0 ft) and an estimated body mass of 10.4 tonnes (23,000 lb). The African bush elephant is characterised by its long prehensile trunk with two finger-like processes; a convex back; large ears which help reduce body heat; and sturdy tusks that are noticeably curved. The skin is grey with scanty hairs, and bending cracks which support thermoregulation by retaining water.


The African bush elephant inhabits a variety of habitats such as forests, grasslands, woodlands, wetlands and agricultural land. It is a mixed herbivore feeding mostly on grasses, creepers, herbs, leaves, and bark. The average adult consumes about 150 kg (330 lb) of vegetation and 230 L (51 imp gal; 61 US gal) of water each day. A social animal, the African bush elephant often travels in herds composed of cows and their offspring. Adult bulls usually live alone or in small bachelor groups. During the mating season, males go through a process called musth; a period of high testosterone levels and heightened aggression. For females, the menstrual cycle lasts three to four months, and gestation around 22 months, the longest of any mammal. 

17-11-1993 TSAVO NAT PARK, KENYA - AFRICAN SAVANNAH ELEPHANT

The African bush elephant (Loxodonta africana) is one of two living African elephant species. It is the largest living terrestrial animal and is distributed across 37 African countries. Since 2021, it has been listed as Endangered on the IUCN Red List. It is threatened foremost by habitat destruction, and in parts of its range also by poaching for meat and ivory.

African bush elephants occur in Sub-Saharan Africa including Uganda, Kenya, Tanzania, Botswana, Zimbabwe, Namibia, Zambia, and Angola. They move between a variety of habitats, including forests, dry and seasonally flooded grasslands, woodlands, shrubland, and wetlands to mountain slopes. In Mali and Namibia, they also inhabit desert and semi-desert areas.

The African bush elephant has grey skin with scanty hairs. Its large ears cover the whole shoulder and can grow as large as 2 m × 1.5 m (6 ft 7 in × 4 ft 11 in). Its large ears help to reduce body heat; flapping them creates air currents and exposes large blood vessels on the inner sides to increase heat loss during hot weather. The African bush elephant's ears are pointed and triangular-shaped. Its occipital plane slopes forward. Its back is shaped markedly concave. Its sturdy tusks are curved out and point forward. The trunk is a prehensile extension of the upper lip and nose. Short tactile hair grows on the trunk, which has two finger-like processes on the tip. This highly sensitive organ is innervated primarily by the trigeminal nerve and is thought to be manipulated by about 40,000-60,000 muscles. Because of this muscular structure, the trunk is so strong that elephants can use it for lifting about 3% of their own body weight. They use it for smelling, touching, feeding, drinking, dusting, sound production, loading, defending, and attacking. Both sexes have tusks, which erupt when they are 1-3 years old and grow throughout life. Tusks grow from deciduous teeth known as tushes that develop in the upper jaw and consist of a crown, root, and pulpal cavity, which are completely formed soon after birth. Tushes reach a length of 5 cm (2.0 in). The tusks of males grow faster than the tusks of females.

Savanna elephants live in a rather complex social hierarchy. These animals gather into family units, consisting of about 10 females and their offspring. Reaching maturity, male calves usually leave the family unit, forming bachelor herds or living solitarily. As a general rule, males socialize with these family groups only when mating. Meanwhile, several family herds may gather together, making up a 'clan'. Each clan is dominated by a female matriarch and can consist of as many as several hundred elephants. African bush elephants are very careful and protective animals. Allomothering is a common practice in this species: females can raise calves of other females of their herd. They protect and care for the calves of the herd, while all adults are sleeping. If a calf strays too far, these allomothers are responsible for retrieving the baby. These active animals are constantly on the move. These elephants forage during the daytime hours, wandering the home range of their herd. Savanna elephants freely communicate with conspecifics both verbally and non-verbally.

17-11-1993 TSAVO NAT PARK, KENYA - BEISA ORYX (Oryx beisa. sspbeisa)


The East African oryx (Oryx beisa), also known as the beisa, is a species of medium-sized antelope from East Africa. It has two subspecies: the common beisa oryx (Oryx beisa beisa) found in steppe and semidesert throughout the Horn of Africa and north of the Tana River, and the fringe-eared oryx (Oryx beisa callotis) south of the Tana River in southern Kenya and parts of Tanzania. The species is listed as Endangered by the IUCN.

In the past, some taxonomists considered it a subspecies of the gemsbok (Oryx gazella), but they are genetically distinct; the diploid chromosome count is 56 for the beisa and 58 for the gemsbok.


The East African oryx stands just over a metre at the shoulder and weighs around 175 lb (79 kg). It has a grey coat with a white underside, separated from the grey by a stripe of black, with black stripes where the head attaches to the neck, along the nose, and from the eye to the mouth and on the forehead. The mane is small and chestnut-coloured; the ringed horns are thin and straight. They are found on both sexes and typically measure 75–80 cm (30–31 in). Comparably, the gemsbok has an entirely black tail, a black patch at the base of the tail, and more black on the legs (including a patch on the hindlegs) and lower flanks. The smaller Arabian oryx is overall whiter with largely dark legs.

East African oryx live in semidesert and steppes, where they eat grasses, leaves, fruit and buds. They are able to store water by raising their body temperatures (so as to avoid perspiration). They gather in herds of five to 40 animals, often with females moving at the front and a large male guarding from the rear. Some older males are solitary. Radio tracking studies show the solitary males are often accompanied for brief periods by breeding-condition females, so it is probable they are executing a strategy to maximise their chances of reproduction.

17-11-1993 TSAVO NAT PARK, KENYA - COMMON IMPALA (FEMALE) (Aepyceros melampus ssp. melampus)


The impala or rooibok (Aepyceros melampus, lit. 'black-footed high-horn' in Ancient Greek) is a medium-sized antelope found in eastern and southern Africa. The only extant member of the genus Aepyceros, and tribe Aepycerotini, it was first described to Europeans by German zoologist Hinrich Lichtenstein in 1812. Two subspecies are recognised—the grassland-dwelling common impala (sometimes referred to as the Kenyan impala), and the larger and darker black-faced impala, which lives in slightly more arid, scrubland environments. The impala reaches 70–92 cm (28–36 in) at the shoulder and weighs 40–76 kg (88–168 lb). It features a glossy, reddish brown coat. The male's slender, lyre-shaped horns are 45–92 cm (18–36 in) long.


Active mainly during the day, the impala may be gregarious or territorial depending upon the climate and geography. Three distinct social groups can be observed: the territorial males, bachelor herds and female herds. The impala is known for two characteristic leaps that constitute an anti-predator strategy. Browsers as well as grazers, impala feed on monocots, dicots, forbs, fruits and acacia pods (whenever available). An annual, three-week-long rut takes place toward the end of the wet season, typically in May. Rutting males fight over dominance, and the victorious male courts females in oestrus. Gestation lasts six to seven months, following which a single calf is born and immediately concealed in cover. Calves are suckled for four to six months; young males—forced out of the all-female groups—join bachelor herds, while females may stay back.

17-11-1993 SERENGETI NAT PARK, KENYA - NUBIAN WOODPECKER (Campethera nubica)


The Nubian woodpecker (Campethera nubica) is a species of bird in the family Picidae. It is distributed widely in Central and Eastern Africa, from Chad in west to Somalia in east and Tanzania in south. It is a fairly common species with a wide range, the population seems stable, and the International Union for Conservation of Nature has rated its conservation status as being of "least concern".

The Nubian woodpecker Is a medium-sized species growing to a length of about 21 cm (8.3 in). The male has a red crown and nape and a reddish streak on the cheek, while the female has a black crown speckled with white, a red nape, and a dark cheek stripe with white speckling. In other respects, the sexes are similar. The upper parts are olive-brown with much cream speckling and barring. The wings are greenish-brown barred with white and the tail greenish-yellow barred with brown, the shafts of the feathers being gold. The throat is cream and the head, neck, breast and belly are white, spotted and barred with black. The beak is grey with a dark tip, the eye red or pink, the orbital ring grey and the feet olive or grey. Various vocalisations are made, shrill ringing or piping notes repeated, often musical but sometimes metallic. Often sung in duet.

The Nubian woodpecker is endemic to eastern Africa. Its range includes Chad, Democratic Republic of the Congo, Djibouti, Eritrea, Ethiopia, Kenya, Somalia, South Sudan, Sudan, Tanzania and Uganda. Its typical habitat is open savannah woodland, especially with Acacia and Euphorbia, bushy areas and scrub. It is a non-migratory bird, and is found at altitudes up to 2,000 m (6,600 ft).Behaviour and ecology

The Nubian woodpecker often feeds alone, keeping in touch with its mate vocally. It forages mostly in trees, but also on the ground, for ants and termites, also consuming spiders and beetles.

Tuesday, 16 November 1993

16-11-1993 SAMBURU, KENYA - GREEN WOOD HOOPOE (Phoeniculus purpureus)


The green wood hoopoe, Phoeniculus purpureus, is a striking bird native to the African continent. It is a large species, reaching up to 44 cm in length, and is adorned with a lustrous metallic dark green plumage. The bird's back is a resplendent purple, and it boasts a remarkably long, diamond-shaped tail of the same hue. Its wings are marked with distinctive white patterns, and the tail edges are decorated with white chevrons, making it a sight to behold.

Upon sighting, one can identify the green wood hoopoe by its long, thin, and curved red bill, which is a prominent feature against its dark plumage. Both sexes appear similar, though the immature birds can be distinguished by their black bills. The white markings on the wings and tail are key identifiers for this species.

The green wood hoopoe is a common resident of sub-Saharan Africa's forests, woodlands, and even suburban gardens, where it can be seen foraging and socializing in groups.

This bird has a wide range across sub-Saharan Africa, with its presence noted from Senegal to South Africa. It is divided into six recognized subspecies, each occupying different geographical regions within the continent.

16-11-1993 SAMBURU, KENYA - SOMALI FISCAL SHRIKE (Lanius somalicus)


The Somali fiscal (Lanius somalicus) is a species of bird in the family Laniidae. Other common names include the Karoli fiscal, the Somali fiscal shrike and the Somali shrike. The bird is found in Djibouti, Ethiopia and Somalia in the Horn of Africa, as well as in Kenya in the African Great Lakes region. Its natural habitat is subtropical or tropical dry shrubland.

The Somali fiscal Is found in East Africa, its range including much of Somalia, southern Djibouti, eastern and southern Ethiopia, southeastern South Sudan and northern Kenya. It is mainly found in open grassy habitats and semi-desert areas with some scrub. Where its range overlaps that of the Taita fiscal (Lanius dorsalis), that bird usually inhabits more densely-vegetated and less arid habitats.

Like other shrikes, the Somali fiscal perches in a position where it has a good view all round and pounces on any suitable prey it observes. It feeds mostly on large insects such as grasshoppers, mantises and beetles. Prey is sometimes impaled on thorns for consumption later, and small birds have sometimes been found in these larders. During the breeding season, it will defend the area around its nest, being prepared to attack any bird that ventures near up to the size of a raven. Breeding usually takes place in the rainy season. The nest is usually in a bush, some 1 to 1.5 m (3 to 5 ft) above the ground, with a clutch of four, or sometimes three, eggs being laid. Little is known of incubation and fledging times.

The Somali fiscal has a wide range and is reported to be common in places. The population trend appears to be steady so the International Union for Conservation of Nature has listed the species as being of "least concern".

16-11-1993 SAMBURU NAT PARK, KENYA - DROMEDARY CAMEL (Camelus dromedarius)


The dromedary (Camelus dromedarius), also known as the dromedary camel, Arabian camel and one-humped camel, is a large camel of the genus Camelus with one hump on its back. It is the tallest of the three camel species; adult males stand 1.8–2.4 m (5 ft 11 in – 7 ft 10 in) at the shoulder, while females are 1.7–1.9 m (5 ft 7 in – 6 ft 3 in) tall. Males typically weigh between 400 and 690 kg (880 and 1,520 lb), and females weigh between 300 and 540 kg (660 and 1,190 lb). The dromedary's distinctive features include its long, curved neck, narrow chest, a single hump, and long hairs on the throat, shoulders, and hump. The coat is generally a shade of brown. The hump, 20 cm (7.9 in) tall or more, is made of fat bound together by fibrous tissue.

The dromedary feeds on foliage and desert vegetation; several adaptations, such as the ability to tolerate losing more than 30% of its total water content, allow it to thrive in its desert habitat. Mating occurs annually and peaks in the rainy season; females bear a single calf after a gestation of 15 months. It is mainly active during daylight hours and forms herds of about 20 individuals, which are led by a dominant male.

The dromedary has not occurred naturally in the wild for nearly 2,000 years. It was probably first domesticated in the Arabian Peninsula about 4,000 years ago, or in Somalia where paintings of it found in Laas Geel date from 5,000 to 9,000 years ago. In the wild, the dromedary inhabited arid regions, including the Sahara. The domesticated dromedary is generally found in the semiarid to arid regions of the Old World, mainly in Africa and the Arabian Peninsula, and a significant feral population occurs in Australia. Products of the dromedary, including its meat and milk, support several North African tribes; it is also commonly used for riding and as a pack animal.

Monday, 15 November 1993

15-11-1993 AMBOSELI, KENYA - SOUTHERN LION (MALE) (Panthera leo ssp. melanochaita)


Panthera leo melanochaita is a lion subspecies in Southern and East Africa. In this part of Africa, lion populations are regionally extinct in Lesotho, Djibouti and Eritrea, and are threatened by loss of habitat and prey base, killing by local people in retaliation for loss of livestock, and in several countries also by trophy hunting. Since the turn of the 21st century, lion populations in intensively managed protected areas in Botswana, Namibia, South Africa and Zimbabwe have increased, but declined in East African range countries. In 2005, a Lion Conservation Strategy was developed for East and Southern Africa.


Results of a phylogeographic study indicate that lion populations in southern and eastern Africa form a major clade distinct from lion populations in West Africa, Central Africa and Asia. In 2017, the Cat Classification Task Force of the IUCN Cat Specialist Group subsumed lion populations according to the major clades into two subspecies, namely P. l. leo and P. l. melanochaita. Within P. l. melanochaita three subclades are clearly distinguishable. One from northeastern Africa, another one from southwestern Africa and a third one from southeastern Africa.


The type specimen for P. l. melanochaita was a black-maned lion from the Cape of Good Hope, known as the Cape lion. Phylogeographic analysis of lion samples from Gabon and the Republic of the Congo indicate their close genetic relation to P. l. melanochaita samples from Namibia and Botswana. It has been referred to as the Southern lion, Southern African lion, East-Southern African lion and the "southern subspecies".

15-11-1993 MASAI MARA, KENYA - COMMON WARTHOG (Phacochoerus Africus)


The common warthog (Phacochoerus africanus) is a wild member of the pig family (Suidae) found in grassland, savanna, and woodland in sub-Saharan Africa. In the past, it was commonly treated as a subspecies of P. aethiopicus, but today that scientific name is restricted to the desert warthog of northern Kenya, Somalia, and eastern Ethiopia.

The common warthog is a medium-sized species, with a head-and-body length ranging from 0.9 to 1.5 m (2 ft 11 in to 4 ft 11 in), and shoulder height from 63.5 to 85 cm (25.0 to 33.5 in). Females, at 45 to 75 kg (99 to 165 lb), are smaller and lighter than males, at 60 to 150 kg (130 to 330 lb).] A warthog is identifiable by the two pairs of tusks protruding from the mouth and curving upwards. The lower pair, which is far shorter than the upper pair, becomes razor-sharp by rubbing against the upper pair every time the mouth is opened and closed. The upper canine teeth can grow to 25.5 cm (10.0 in) long and have a wide elliptical cross section, being about 4.5 cm (1.8 in) deep and 2.5 cm (0.98 in) wide. A tusk will curve 90° or more from the root, and will not lie flat on a table, as it curves somewhat backwards as it grows. The tusks are not used for digging, but are used for combat with other hogs and in defense against predators – the lower set can inflict severe wounds.

Common warthog ivory is taken from the constantly growing canine teeth. The tusks, particularly the upper set, work in much the same way as elephant tusks with all designs scaled down. Tusks are carved predominantly for the tourist trade in eastern and southern Africa.

15-11-1993 AMBOSELI, KENYA - EASTERN BLACK RHINOCEROS (Diceros bicornis ssp. michaeli)

The eastern black rhinoceros (Diceros bicornis michaeli), also known as the East African black rhinoceros, is a subspecies of the black rhinoceros. Its numbers are very low due to poaching for its horn, and it is listed as critically endangered.

The eastern black rhino is distinguishable from the southern subspecies as it has a longer, leaner, and more curved horn. Its skin is also very grooved. Diceros bicornis michaeli is also reportedly more aggressive than the other three subspecies of black rhino. They are browsers and are usually found in highland forest and savanna habitat.

Once located in Ethiopia, Somalia, Tanzania and Kenya, as of 2017 they can only be found in Kenya (594 animals), Rwanda and in northern Tanzania (80 animals). A population of currently 60 animals is kept outside its natural range in South Africa (Addo Elephant National Park). The population has declined 90% in the last three generations. In 2010 their total numbers were estimated at 740 animals, with an increasing trend. They are threatened mainly from illegal poaching for their horns.

The IUCN figures for Diceros bicornis michaeli also include those for black rhinos from South Sudan, Uganda, southwestern Ethiopia and western Kenya. These are referred to a separate subspecies (Diceros bicornis ladoensis) by some authorities. As the black rhinoceros population is extirpated in most of these areas, the status of the latter subspecies is unclear. Some animals of the Kenyan population may belong to it.

These black rhinos were examined by Benson and others through the Kenya Wildlife Service and had an article published by the African Journal of Ecology (Benson, 1, 791). Each individual rhino's relationship concerning “density dependence...sex ratio, and underlying growth rates” was the team's main focus. The rhino began increasing its population growth to the point when growth rate was nonexistent.

Eastern black rhinos in general are currently listed as threatened under the Conservation Status for a number of factors, but mainly due to illegal poaching. The 90% decline in their population is a hazard as mortality rates increase, as is consistent emigration (Primack, 2002). The reason why the rhino increases its densities and their birth rate decreases is because of the continuous threat of poaching and emigration.

Sex ratio brings about reproduction issues because the male numbers begin to dominate, leaving too low of a number of females to allow positive, consistent birth and growth rates (Benson, 2, 792).

15-8-1993 SERENGETTI, KENYA - RUPPELL'S STARLING (Lamprotornis purpuroptera)


Rüppell's starling (Lamprotornis purpuroptera), also known as Rueppell's glossy-starling or Rueppell's long-tailed starling, is a species of starling in the family Sturnidae. It is found in Burundi, the Democratic Republic of the Congo, Eritrea, Ethiopia, Kenya, Rwanda, Somalia, South Sudan, Sudan, Chad, Tanzania, and Uganda.

15-11-1993 MASAI MARA, KENYA - AFRICAN BUFFALO (Syncerus caffer)


The African buffalo (Syncerus caffer) is a large sub-Saharan African bovine. There are five subspecies that are recognized as being valid. Syncerus caffer caffer, the Cape buffalo, is the nominotypical subspecies, and the largest one, found in Southern and East Africa. S. c. nanus (the forest buffalo) is the smallest subspecies, common in forest areas of Central and West Africa, while S. c. brachyceros is in West Africa and S. c. aequinoctialis is in the savannas of East Africa. The adult African buffalo's horns are its characteristic feature: they have fused bases, forming a continuous bone shield across the top of the head referred to as a "boss". It is widely regarded as one of the most dangerous animals on the African continent, and according to some estimates it gores, tramples, and kills over 200 people every year.


The adult African buffalo's horns are its characteristic feature: they have fused bases that form a continuous bone shield, referred to as a "boss", across the top of the head.

The African buffalo is more closely related to other buffalo species than it is to other bovids such as American bison or domestic cattle, with its closest living relative being the Asian water buffalo. Its unpredictable temperament may be part of the reason that the African buffalo has never been domesticated and has no domesticated descendants, unlike the wild yak and wild water buffalo, which are the ancestors of the domestic yak and water buffalo, respectively. Natural predators of adult African buffaloes include lions, African wild dogs, spotted hyenas, and Nile crocodiles. As one of the Big Five game animals, the Cape buffalo is a sought-after trophy in hunting.

15-11-1993 AMBOSELI, KENYA - SOUTHERN LION (MALE) (Panthera leo ssp. melanochaita)


Panthera leo melanochaita is a lion subspecies in Southern and East Africa. In this part of Africa, lion populations are regionally extinct in Lesotho, Djibouti and Eritrea, and are threatened by loss of habitat and prey base, killing by local people in retaliation for loss of livestock, and in several countries also by trophy hunting. Since the turn of the 21st century, lion populations in intensively managed protected areas in Botswana, Namibia, South Africa and Zimbabwe have increased, but declined in East African range countries. In 2005, a Lion Conservation Strategy was developed for East and Southern Africa.


Results of a phylogeographic study indicate that lion populations in southern and eastern Africa form a major clade distinct from lion populations in West Africa, Central Africa and Asia. In 2017, the Cat Classification Task Force of the IUCN Cat Specialist Group subsumed lion populations according to the major clades into two subspecies, namely P. l. leo and P. l. melanochaita. Within P. l. melanochaita three subclades are clearly distinguishable. One from northeastern Africa, another one from southwestern Africa and a third one from southeastern Africa.


The type specimen for P. l. melanochaita was a black-maned lion from the Cape of Good Hope, known as the Cape lion. Phylogeographic analysis of lion samples from Gabon and the Republic of the Congo indicate their close genetic relation to P. l. melanochaita samples from Namibia and Botswana. It has been referred to as the Southern lion, Southern African lion, East-Southern African lion and the "southern subspecies".

15-11-1993 AMBOSELI, KENYA - THOMSON'S GAZELLE (Eudorcas thomsonii)


Thomson's gazelle (Eudorcas thomsonii) is one of the best known species of gazelles. It is named after explorer Joseph Thomson and is sometimes referred to as a "tommie". It is considered by some to be a subspecies of the red-fronted gazelle and was formerly considered a member of the genus Gazella within the subgenus Eudorcas, before Eudorcas was elevated to genus status.

Thomson's gazelles can be found in numbers exceeding 200,000 in Africa and are recognized as the most common type of gazelle in East Africa. A small fast antelope, the Thomson's gazelle is said to have top speeds up to 80–90 km/h (50–55 mph). It is the fourth-fastest land animal, after the cheetah (its main predator), pronghorn, and springbok.

15-11-1993 AMBOSELI, KENYA - SOUTHERN LION (FEMALE) (Panthera leo ssp. melanochaita)


Panthera leo melanochaita is a lion subspecies in Southern and East Africa. In this part of Africa, lion populations are regionally extinct in Lesotho, Djibouti and Eritrea, and are threatened by loss of habitat and prey base, killing by local people in retaliation for loss of livestock, and in several countries also by trophy hunting. Since the turn of the 21st century, lion populations in intensively managed protected areas in Botswana, Namibia, South Africa and Zimbabwe have increased, but declined in East African range countries. In 2005, a Lion Conservation Strategy was developed for East and Southern Africa.


Results of a phylogeographic study indicate that lion populations in southern and eastern Africa form a major clade distinct from lion populations in West Africa, Central Africa and Asia. In 2017, the Cat Classification Task Force of the IUCN Cat Specialist Group subsumed lion populations according to the major clades into two subspecies, namely P. l. leo and P. l. melanochaita. Within P. l. melanochaita three subclades are clearly distinguishable. One from northeastern Africa, another one from southwestern Africa and a third one from southeastern Africa.


The type specimen for P. l. melanochaita was a black-maned lion from the Cape of Good Hope, known as the Cape lion. Phylogeographic analysis of lion samples from Gabon and the Republic of the Congo indicate their close genetic relation to P. l. melanochaita samples from Namibia and Botswana. It has been referred to as the Southern lion, Southern African lion, East-Southern African lion and the "southern subspecies".

15-11-1993 AMBOSELI, KENYA - SOUTHERN LION (FEMALE) (Panthera leo ssp. melanochaita)


The Southern lion is a lion subspecies from Southern and East Africa. In this region, lion populations were referred to by several regional names, including Katanga lion, Transvaal lion, Kalahari lion, Southeast African lion, Southwest African lion, Masai lion, Serengeti lion, Tsavo lion, and Uganda lion. It has also been referred to as 'Eastern-Southern African lion', 'Southern lion', and as 'southern subspecies'.


The fur of Southern lions varies in color from light buff to dark brown. They have rounded ears and a black tail tuft. Mane color varies from sandy, tawny, isabelline, and light reddish yellow to dark brown and black. Mane length varies from short to extending to knee joints and under the belly. Lions without a mane were observed in the Tsavo area. Mane development is related to age: older males have more extensive manes than younger ones; manes continue to grow up to the age of four to five years, long after lions become reproductively mature. 


Males living in the Kenyan highlands develop heavier manes than lions in the more humid and warmer lowlands of eastern and northern Kenya. White lions have occasionally been encountered in and around South Africa's Kruger National Park and the adjacent Timbavati Private Game Reserve. Their whitish fur is a rare morph caused by a double recessive allele. It has normal pigmentation in the eyes and skin. They were removed from the wild in the 1970s, thus decreasing the white lion gene pool.

15-11-1993 AMBOSELI, KENYA - OLIVE BABOON (Papio anubi)





The olive baboon (Papio anubis), also called the Anubis baboon, is a member of the family Cercopithecidae Old World monkeys. The species is the most wide-ranging of all baboons, being native to 25 countries throughout Africa, extending from Mali eastward to Ethiopia and Tanzania. Isolated populations are also present in some mountainous regions of the Sahara. It inhabits savannahs, steppes, and forests. The common name is derived from its coat colour, which is a shade of green-grey at a distance. A variety of communications, vocal and non-vocal, facilitate a complex social structure.

The olive baboon is named for its coat, which, at a distance, is a shade of green-grey. At closer range, its coat is multicoloured, due to rings of yellow-brown and black on the hairs. The hair on the baboon's face is coarser and ranges from dark grey to black. This coloration is shared by both sexes, although males have a mane of longer hair that tapers down to ordinary length along the back.


Besides the mane, the male olive baboon differs from the female in terms of weight, body and canine tooth size; males are, on average, 70 cm (28 in) tall while standing and females measure 60 cm (24 in) in height. The olive baboon is one of the largest species of monkey; along with chacma baboons, mandrills, and muriquis. The head-and-body length can range from 50 to 114 cm (20 to 45 in), with a species average of around 85 cm (33 in). At the shoulder on all fours, females average 55 cm (22 in) against males, which average 70 cm (28 in). The typical weight range for both sexes is reportedly 10–37 kg (22–82 lb), with males averaging 24 kg (53 lb) and females averaging 14.7 kg (32 lb). Some males may weigh as much as 50 kg (110 lb).

Like other baboons, the olive baboon has an elongated, dog-like muzzle. Its 38 to 58 cm (15 to 23 in) long tail and four-legged gait can make it seem canine. The tail almost looks as if it is broken, as it is erect for the first quarter, after which it drops down sharply. The bare patch of a baboon's rump is smaller in the olive baboon than in the Hamadryas baboon or Guinea baboon. The olive baboon, like most cercopithecines, has a cheek pouch with which to store food.


The species inhabits a strip of 25 equatorial African countries, very nearly ranging from the east to west coasts of the continent. The exact boundaries of this strip are not clearly defined, as the species' territory overlaps with that of other baboon species. In many places, this has resulted in cross-breeding between species. For example, considerable hybridisation has occurred between the olive baboon and the hamadryas baboon in Ethiopia. Cross-breeding with the yellow baboon and the Guinea baboon has also been observed. Although this has been noted, the hybrids have not as yet been well studied.

Throughout its wide range, the olive baboon can be found in a number of different habitats. It is usually classified as savannah-dwelling, living in the wide plains of the grasslands. The grasslands, especially those near open woodland, do make up a large part of its habitat, but the baboon also inhabits rainforests and deserts. Uganda and the Democratic Republic of the Congo, for instance, both support olive baboon populations in dense tropical forests.

15-11-1993 AMBOSELI, KENYA - EASTERN BLACK RHINOCEROS (Diceros bicornis ssp. michaeli)


The eastern black rhinoceros (Diceros bicornis michaeli), also known as the East African black rhinoceros, is a subspecies of the black rhinoceros. Its numbers are very low due to poaching for its horn, and it is listed as critically endangered.

The eastern black rhino is distinguishable from the southern subspecies as it has a longer, leaner, and more curved horn. Its skin is also very grooved. Diceros bicornis michaeli is also reportedly more aggressive than the other three subspecies of black rhino. They are browsers and are usually found in highland forest and savanna habitat.

Once located in Ethiopia, Somalia, Tanzania and Kenya, as of 2017 they can only be found in Kenya (594 animals), Rwanda and in northern Tanzania (80 animals). A population of currently 60 animals is kept outside its natural range in South Africa (Addo Elephant National Park). The population has declined 90% in the last three generations. In 2010 their total numbers were estimated at 740 animals, with an increasing trend. They are threatened mainly from illegal poaching for their horns.

The IUCN figures for Diceros bicornis michaeli also include those for black rhinos from South Sudan, Uganda, southwestern Ethiopia and western Kenya. These are referred to a separate subspecies (Diceros bicornis ladoensis) by some authorities. As the black rhinoceros population is extirpated in most of these areas, the status of the latter subspecies is unclear. Some animals of the Kenyan population may belong to it.

These black rhinos were examined by Benson and others through the Kenya Wildlife Service and had an article published by the African Journal of Ecology (Benson, 1, 791). Each individual rhino's relationship concerning “density dependence...sex ratio, and underlying growth rates” was the team's main focus. The rhino began increasing its population growth to the point when growth rate was nonexistent.

15-11-1993 AMBOSELI, KENYA - AFRICAN ELEPHANT (Loxodonta africana)


The African bush elephant (Loxodonta africana) is one of two living African elephant species. It is the largest living terrestrial animal and is distributed across 37 African countries. Since 2021, it has been listed as Endangered on the IUCN Red List. It is threatened foremost by habitat destruction, and in parts of its range also by poaching for meat and ivory.

The African bush elephant has grey skin with scanty hairs. Its large ears cover the whole shoulder and can grow as large as 2 m × 1.5 m (6 ft 7 in × 4 ft 11 in). Its large ears help to reduce body heat; flapping them creates air currents and exposes large blood vessels on the inner sides to increase heat loss during hot weather. The African bush elephant's ears are pointed and triangular-shaped. Its occipital plane slopes forward. Its back is shaped markedly concave. Its sturdy tusks are curved out and point forward. The trunk is a prehensile extension of the upper lip and nose. Short tactile hair grows on the trunk, which has two finger-like processes on the tip. This highly sensitive organ is innervated primarily by the trigeminal nerve and is thought to be manipulated by about 40,000-60,000 muscles. Because of this muscular structure, the trunk is so strong that elephants can use it for lifting about 3% of their own body weight. They use it for smelling, touching, feeding, drinking, dusting, sound production, loading, defending, and attacking. Both sexes have tusks, which erupt when they are 1-3 years old and grow throughout life. Tusks grow from deciduous teeth known as tushes that develop in the upper jaw and consist of a crown, root, and pulpal cavity, which are completely formed soon after birth. Tushes reach a length of 5 cm (2.0 in). The tusks of males grow faster than the tusks of females.


African bush elephants occur in Sub-Saharan Africa including Uganda, Kenya, Tanzania, Botswana, Zimbabwe, Namibia, Zambia, and Angola. They move between a variety of habitats, including forests, dry and seasonally flooded grasslands, woodlands, shrubland, and wetlands to mountain slopes. In Mali and Namibia, they also inhabit desert and semi-desert areas.

Savanna elephants live in a rather complex social hierarchy. These animals gather into family units, consisting of about 10 females and their offspring. Reaching maturity, male calves usually leave the family unit, forming bachelor herds or living solitarily. As a general rule, males socialize with these family groups only when mating. Meanwhile, several family herds may gather together, making up a 'clan'. Each clan is dominated by a female matriarch and can consist of as many as several hundred elephants. African bush elephants are very careful and protective animals. Allomothering is a common practice in this species: females can raise calves of other females of their herd. They protect and care for the calves of the herd, while all adults are sleeping. If a calf strays too far, these allomothers are responsible for retrieving the baby. These active animals are constantly on the move. These elephants forage during the daytime hours, wandering the home range of their herd. Savanna elephants freely communicate with conspecifics both verbally and non-verbally.