The meadow brown (Maniola jurtina) is a butterfly found in the Palearctic realm. Its range includes Europe south of 62°N, Russia eastwards to the Urals, Asia Minor, Iraq, Iran, North Africa and the Canary Islands. The larvae feed on grasses.
There is marked sexual dimorphism in this species. The upperside of the male is uniformly light brown with a black ocellus centered white at the apex of the forewing, while the female has a tawny patch more or less extended around this ocella. The underside forewing is ochre-colored bordered with dark beige with the same ocelli at the apex in the male, while the hindwing is greyish to brown with a more or less orange band in the female. The males are also much more active and range far about, while females fly less and often may not move away from the area where they grew up.
A variable number of smaller eyespots are usually found on the hindwing undersides. These may number up to twelve per butterfly, with up to six on each wing, but sometimes none. The factors that govern polymorphism in this trait are not resolved, although a number of theories have been proposed (Stevens 2005). On the other hand, the evolutionary significance of the upperwing eyespots is more obvious: The more active males have a markedly more cryptic upperside pattern, whereas the females have more opportunity to present their eyespots in a sudden display of colors and patterns that presumably startle predators so the butterfly has a better chance of escaping. Some specimens are bi-pupilled.
